LeishMANIAdb
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Sel1 repeat family protein

Quick info Annotations Function or PPIs Localization Expansion Sequence features Structure Function Putative motif mimicry Homologs Download

Quick info

Protein:
Sel1 repeat family protein
Gene product:
Sel1 repeat, putative
Species:
Leishmania major
UniProt:
Q4QIN3_LEIMA
TriTrypDb:
LmjF.07.0590 * , LMJLV39_070012100 * , LMJSD75_070012300 *
Length:
1257

Annotations

LeishMANIAdb annotations

Fast evolving protein with mostly disordered structure. Possibly coat proteins.

Localization

Secreted promastigote
Source Evidence on protein Close homologs
Cuervo et al. no yes: 0
Hassani et al. no yes: 0
Forrest at al. (metacyclic) no yes: 0
Forrest at al. (procyclic) no yes: 0
Silverman et al. no yes: 0
Pissara et al. no yes: 0
Secreted amastigote
Source Evidence on protein Close homologs
Pires et al. no yes: 0
Exosome
Source Evidence on protein Close homologs
Silverman et al. no yes: 0
Glycosome
Source Evidence on protein Close homologs
Jamdhade et al. no yes: 0
Predictions
Source Evidence on protein Close homologs
DeepLoc
SignalP6 no yes: 2, no: 4
NetGPI no yes: 0, no: 6
Cellular components
Term Name Level Count
GO:0005789 endoplasmic reticulum membrane 4 2
GO:0016020 membrane 2 4
GO:0031090 organelle membrane 3 2
GO:0110165 cellular anatomical entity 1 4

Expansion

Sequence features

Q4QIN3
Sequence
MSA
Disorder
Secondary
Topology
Domains
SignalP
GPI
Phosphorylations
ELMs

Structure

Predicted structure by AlphaFold2

Related structures:

AlphaFold database: Q4QIN3

Function

Biological processes
Term Name Level Count
GO:0006508 proteolysis 4 2
GO:0006807 nitrogen compound metabolic process 2 2
GO:0006950 response to stress 2 2
GO:0008152 metabolic process 1 2
GO:0009056 catabolic process 2 2
GO:0009057 macromolecule catabolic process 4 2
GO:0009987 cellular process 1 2
GO:0010033 response to organic substance 3 2
GO:0010243 response to organonitrogen compound 4 2
GO:0010498 proteasomal protein catabolic process 5 2
GO:0019538 protein metabolic process 3 2
GO:0030163 protein catabolic process 4 2
GO:0033554 cellular response to stress 3 2
GO:0034976 response to endoplasmic reticulum stress 4 2
GO:0036503 ERAD pathway 5 2
GO:0042221 response to chemical 2 2
GO:0043170 macromolecule metabolic process 3 2
GO:0044238 primary metabolic process 2 2
GO:0050896 response to stimulus 1 2
GO:0051603 proteolysis involved in protein catabolic process 5 2
GO:0051716 cellular response to stimulus 2 2
GO:0071704 organic substance metabolic process 2 2
GO:1901564 organonitrogen compound metabolic process 3 2
GO:1901565 organonitrogen compound catabolic process 4 2
GO:1901575 organic substance catabolic process 3 2
GO:1901698 response to nitrogen compound 3 2
Could not find GO molecular_function term for this entry.

Putative motif mimicry

Leishmania From To Domain/Motif Score
CLV_C14_Caspase3-7 253 257 PF00656 0.570
CLV_C14_Caspase3-7 421 425 PF00656 0.403
CLV_C14_Caspase3-7 467 471 PF00656 0.346
CLV_C14_Caspase3-7 644 648 PF00656 0.536
CLV_C14_Caspase3-7 819 823 PF00656 0.636
CLV_C14_Caspase3-7 831 835 PF00656 0.503
CLV_C14_Caspase3-7 888 892 PF00656 0.513
CLV_NRD_NRD_1 219 221 PF00675 0.772
CLV_NRD_NRD_1 28 30 PF00675 0.702
CLV_NRD_NRD_1 522 524 PF00675 0.742
CLV_NRD_NRD_1 526 528 PF00675 0.680
CLV_NRD_NRD_1 564 566 PF00675 0.558
CLV_NRD_NRD_1 605 607 PF00675 0.618
CLV_NRD_NRD_1 700 702 PF00675 0.618
CLV_NRD_NRD_1 807 809 PF00675 0.860
CLV_PCSK_FUR_1 805 809 PF00082 0.788
CLV_PCSK_KEX2_1 1194 1196 PF00082 0.749
CLV_PCSK_KEX2_1 219 221 PF00082 0.772
CLV_PCSK_KEX2_1 526 528 PF00082 0.727
CLV_PCSK_KEX2_1 564 566 PF00082 0.558
CLV_PCSK_KEX2_1 605 607 PF00082 0.618
CLV_PCSK_KEX2_1 700 702 PF00082 0.618
CLV_PCSK_KEX2_1 807 809 PF00082 0.871
CLV_PCSK_PC1ET2_1 1194 1196 PF00082 0.749
CLV_PCSK_SKI1_1 1195 1199 PF00082 0.718
CLV_PCSK_SKI1_1 366 370 PF00082 0.643
CLV_PCSK_SKI1_1 423 427 PF00082 0.542
CLV_PCSK_SKI1_1 605 609 PF00082 0.618
CLV_PCSK_SKI1_1 700 704 PF00082 0.578
CLV_PCSK_SKI1_1 705 709 PF00082 0.592
CLV_PCSK_SKI1_1 958 962 PF00082 0.723
CLV_Separin_Metazoa 1047 1051 PF03568 0.559
DEG_APCC_DBOX_1 1194 1202 PF00400 0.418
DEG_APCC_DBOX_1 365 373 PF00400 0.436
DEG_APCC_DBOX_1 526 534 PF00400 0.507
DEG_COP1_1 95 102 PF00400 0.595
DEG_Kelch_Keap1_1 235 240 PF01344 0.575
DEG_SPOP_SBC_1 340 344 PF00917 0.553
DEG_SPOP_SBC_1 872 876 PF00917 0.565
DEG_SPOP_SBC_1 94 98 PF00917 0.500
DOC_ANK_TNKS_1 219 226 PF00023 0.574
DOC_CDC14_PxL_1 1142 1150 PF14671 0.413
DOC_CYCLIN_RxL_1 523 532 PF00134 0.521
DOC_CYCLIN_RxL_1 696 704 PF00134 0.418
DOC_CYCLIN_RxL_1 736 747 PF00134 0.498
DOC_MAPK_gen_1 447 454 PF00069 0.346
DOC_MAPK_gen_1 501 510 PF00069 0.523
DOC_MAPK_gen_1 523 533 PF00069 0.525
DOC_MAPK_MEF2A_6 1118 1125 PF00069 0.521
DOC_MAPK_MEF2A_6 447 454 PF00069 0.418
DOC_MAPK_NFAT4_5 447 455 PF00069 0.418
DOC_MAPK_RevD_3 205 220 PF00069 0.490
DOC_PP1_RVXF_1 401 407 PF00149 0.389
DOC_PP2B_LxvP_1 357 360 PF13499 0.530
DOC_PP4_FxxP_1 836 839 PF00568 0.626
DOC_USP7_MATH_1 1158 1162 PF00917 0.602
DOC_USP7_MATH_1 128 132 PF00917 0.574
DOC_USP7_MATH_1 146 150 PF00917 0.444
DOC_USP7_MATH_1 163 167 PF00917 0.529
DOC_USP7_MATH_1 184 188 PF00917 0.591
DOC_USP7_MATH_1 232 236 PF00917 0.638
DOC_USP7_MATH_1 272 276 PF00917 0.618
DOC_USP7_MATH_1 278 282 PF00917 0.560
DOC_USP7_MATH_1 341 345 PF00917 0.550
DOC_USP7_MATH_1 480 484 PF00917 0.600
DOC_USP7_MATH_1 517 521 PF00917 0.556
DOC_USP7_MATH_1 892 896 PF00917 0.527
DOC_USP7_MATH_1 919 923 PF00917 0.579
DOC_USP7_MATH_1 93 97 PF00917 0.570
DOC_USP7_MATH_1 930 934 PF00917 0.525
DOC_USP7_MATH_1 946 950 PF00917 0.583
DOC_WW_Pin1_4 1071 1076 PF00397 0.552
DOC_WW_Pin1_4 1150 1155 PF00397 0.563
DOC_WW_Pin1_4 197 202 PF00397 0.573
DOC_WW_Pin1_4 358 363 PF00397 0.577
DOC_WW_Pin1_4 594 599 PF00397 0.418
DOC_WW_Pin1_4 908 913 PF00397 0.587
DOC_WW_Pin1_4 983 988 PF00397 0.418
LIG_14-3-3_CanoR_1 1050 1056 PF00244 0.499
LIG_14-3-3_CanoR_1 1090 1096 PF00244 0.594
LIG_14-3-3_CanoR_1 366 372 PF00244 0.439
LIG_14-3-3_CanoR_1 746 751 PF00244 0.528
LIG_14-3-3_CanoR_1 838 847 PF00244 0.532
LIG_BIR_III_2 256 260 PF00653 0.533
LIG_BIR_III_4 1222 1226 PF00653 0.530
LIG_BRCT_BRCA1_1 1073 1077 PF00533 0.561
LIG_BRCT_BRCA1_1 506 510 PF00533 0.599
LIG_CaM_IQ_9 1129 1145 PF13499 0.513
LIG_eIF4E_1 680 686 PF01652 0.418
LIG_FHA_1 1029 1035 PF00498 0.604
LIG_FHA_1 1108 1114 PF00498 0.496
LIG_FHA_1 1185 1191 PF00498 0.445
LIG_FHA_1 1204 1210 PF00498 0.487
LIG_FHA_1 181 187 PF00498 0.536
LIG_FHA_1 291 297 PF00498 0.573
LIG_FHA_1 385 391 PF00498 0.425
LIG_FHA_1 872 878 PF00498 0.606
LIG_FHA_2 1180 1186 PF00498 0.528
LIG_FHA_2 133 139 PF00498 0.644
LIG_FHA_2 165 171 PF00498 0.656
LIG_FHA_2 232 238 PF00498 0.570
LIG_FHA_2 642 648 PF00498 0.530
LIG_FHA_2 86 92 PF00498 0.533
LIG_Integrin_RGD_1 254 256 PF01839 0.813
LIG_KLC1_Yacidic_2 378 383 PF13176 0.422
LIG_LIR_Apic_2 280 286 PF02991 0.628
LIG_LIR_Apic_2 834 839 PF02991 0.501
LIG_LIR_Gen_1 430 438 PF02991 0.346
LIG_LIR_Gen_1 547 557 PF02991 0.418
LIG_LIR_Gen_1 842 853 PF02991 0.502
LIG_LIR_Gen_1 879 887 PF02991 0.497
LIG_LIR_Gen_1 895 902 PF02991 0.492
LIG_LIR_Gen_1 903 913 PF02991 0.513
LIG_LIR_Gen_1 950 961 PF02991 0.540
LIG_LIR_Nem_3 347 352 PF02991 0.481
LIG_LIR_Nem_3 370 374 PF02991 0.425
LIG_LIR_Nem_3 507 513 PF02991 0.597
LIG_LIR_Nem_3 547 553 PF02991 0.418
LIG_LIR_Nem_3 556 560 PF02991 0.312
LIG_LIR_Nem_3 842 848 PF02991 0.504
LIG_LIR_Nem_3 879 883 PF02991 0.504
LIG_LIR_Nem_3 895 899 PF02991 0.492
LIG_LIR_Nem_3 903 908 PF02991 0.500
LIG_LIR_Nem_3 950 956 PF02991 0.538
LIG_MYND_1 598 602 PF01753 0.418
LIG_MYND_1 926 930 PF01753 0.498
LIG_MYND_3 1145 1149 PF01753 0.443
LIG_NRBOX 1197 1203 PF00104 0.410
LIG_NRBOX 403 409 PF00104 0.380
LIG_NRBOX 547 553 PF00104 0.389
LIG_NRBOX 848 854 PF00104 0.415
LIG_NRBOX 951 957 PF00104 0.493
LIG_PCNA_yPIPBox_3 708 722 PF02747 0.412
LIG_PCNA_yPIPBox_3 840 853 PF02747 0.424
LIG_Pex14_2 1014 1018 PF04695 0.453
LIG_PTB_Apo_2 1012 1019 PF02174 0.443
LIG_PTB_Apo_2 976 983 PF02174 0.418
LIG_PTB_Phospho_1 976 982 PF10480 0.418
LIG_REV1ctd_RIR_1 508 518 PF16727 0.590
LIG_SH2_CRK 1240 1244 PF00017 0.411
LIG_SH2_CRK 244 248 PF00017 0.607
LIG_SH2_CRK 557 561 PF00017 0.374
LIG_SH2_CRK 845 849 PF00017 0.537
LIG_SH2_CRK 953 957 PF00017 0.562
LIG_SH2_NCK_1 1052 1056 PF00017 0.607
LIG_SH2_NCK_1 244 248 PF00017 0.578
LIG_SH2_SRC 905 908 PF00017 0.532
LIG_SH2_STAP1 1109 1113 PF00017 0.493
LIG_SH2_STAP1 180 184 PF00017 0.569
LIG_SH2_STAP1 352 356 PF00017 0.546
LIG_SH2_STAP1 429 433 PF00017 0.346
LIG_SH2_STAT3 634 637 PF00017 0.492
LIG_SH2_STAT5 1052 1055 PF00017 0.605
LIG_SH2_STAT5 1070 1073 PF00017 0.385
LIG_SH2_STAT5 1103 1106 PF00017 0.507
LIG_SH2_STAT5 1109 1112 PF00017 0.471
LIG_SH2_STAT5 371 374 PF00017 0.425
LIG_SH2_STAT5 381 384 PF00017 0.469
LIG_SH2_STAT5 550 553 PF00017 0.346
LIG_SH2_STAT5 569 572 PF00017 0.346
LIG_SH2_STAT5 634 637 PF00017 0.420
LIG_SH2_STAT5 669 672 PF00017 0.471
LIG_SH2_STAT5 901 904 PF00017 0.509
LIG_SH2_STAT5 905 908 PF00017 0.518
LIG_SH2_STAT5 941 944 PF00017 0.592
LIG_SH2_STAT5 974 977 PF00017 0.418
LIG_SH3_3 353 359 PF00018 0.578
LIG_SH3_3 596 602 PF00018 0.418
LIG_SUMO_SIM_anti_2 1244 1249 PF11976 0.408
LIG_SUMO_SIM_anti_2 879 885 PF11976 0.501
LIG_SUMO_SIM_par_1 882 889 PF11976 0.523
LIG_TRAF2_1 1038 1041 PF00917 0.591
LIG_TRAF2_1 149 152 PF00917 0.623
LIG_TRAF2_1 167 170 PF00917 0.544
LIG_TRAF2_1 234 237 PF00917 0.647
LIG_TRAF2_1 267 270 PF00917 0.596
LIG_TRAF2_1 47 50 PF00917 0.573
LIG_TRAF2_1 858 861 PF00917 0.526
LIG_TRFH_1 1142 1146 PF08558 0.493
LIG_TYR_ITIM 1238 1243 PF00017 0.394
LIG_TYR_ITIM 372 377 PF00017 0.508
LIG_TYR_ITIM 548 553 PF00017 0.411
LIG_TYR_ITIM 951 956 PF00017 0.718
LIG_TYR_ITIM 997 1002 PF00017 0.512
LIG_UBA3_1 530 534 PF00899 0.611
LIG_WRC_WIRS_1 279 284 PF05994 0.612
LIG_WRC_WIRS_1 428 433 PF05994 0.411
LIG_WRC_WIRS_1 877 882 PF05994 0.648
LIG_WW_1 1067 1070 PF00397 0.766
MOD_CK1_1 1028 1034 PF00069 0.534
MOD_CK1_1 1150 1156 PF00069 0.670
MOD_CK1_1 1161 1167 PF00069 0.678
MOD_CK1_1 1203 1209 PF00069 0.663
MOD_CK1_1 329 335 PF00069 0.802
MOD_CK1_1 339 345 PF00069 0.669
MOD_CK1_1 457 463 PF00069 0.512
MOD_CK1_1 53 59 PF00069 0.744
MOD_CK1_1 723 729 PF00069 0.668
MOD_CK1_1 749 755 PF00069 0.637
MOD_CK1_1 801 807 PF00069 0.666
MOD_CK1_1 862 868 PF00069 0.788
MOD_CK1_1 876 882 PF00069 0.544
MOD_CK2_1 1035 1041 PF00069 0.768
MOD_CK2_1 1164 1170 PF00069 0.802
MOD_CK2_1 1179 1185 PF00069 0.650
MOD_CK2_1 146 152 PF00069 0.787
MOD_CK2_1 163 169 PF00069 0.637
MOD_CK2_1 170 176 PF00069 0.752
MOD_CK2_1 231 237 PF00069 0.831
MOD_CK2_1 264 270 PF00069 0.764
MOD_CK2_1 278 284 PF00069 0.579
MOD_CK2_1 68 74 PF00069 0.706
MOD_CK2_1 812 818 PF00069 0.781
MOD_CK2_1 838 844 PF00069 0.761
MOD_CK2_1 85 91 PF00069 0.615
MOD_CK2_1 862 868 PF00069 0.746
MOD_CK2_1 876 882 PF00069 0.544
MOD_CK2_1 95 101 PF00069 0.780
MOD_Cter_Amidation 562 565 PF01082 0.411
MOD_GlcNHglycan 1037 1040 PF01048 0.758
MOD_GlcNHglycan 1053 1056 PF01048 0.758
MOD_GlcNHglycan 1091 1094 PF01048 0.769
MOD_GlcNHglycan 1149 1153 PF01048 0.671
MOD_GlcNHglycan 1158 1161 PF01048 0.782
MOD_GlcNHglycan 1163 1166 PF01048 0.643
MOD_GlcNHglycan 130 133 PF01048 0.745
MOD_GlcNHglycan 172 175 PF01048 0.781
MOD_GlcNHglycan 334 337 PF01048 0.735
MOD_GlcNHglycan 338 341 PF01048 0.702
MOD_GlcNHglycan 459 462 PF01048 0.512
MOD_GlcNHglycan 482 485 PF01048 0.718
MOD_GlcNHglycan 498 501 PF01048 0.613
MOD_GlcNHglycan 504 507 PF01048 0.723
MOD_GlcNHglycan 577 580 PF01048 0.568
MOD_GlcNHglycan 641 644 PF01048 0.695
MOD_GlcNHglycan 814 817 PF01048 0.768
MOD_GlcNHglycan 849 852 PF01048 0.618
MOD_GlcNHglycan 896 899 PF01048 0.531
MOD_GlcNHglycan 921 924 PF01048 0.708
MOD_GlcNHglycan 932 935 PF01048 0.678
MOD_GlcNHglycan 942 945 PF01048 0.673
MOD_GSK3_1 1150 1157 PF00069 0.663
MOD_GSK3_1 1200 1207 PF00069 0.647
MOD_GSK3_1 128 135 PF00069 0.742
MOD_GSK3_1 176 183 PF00069 0.852
MOD_GSK3_1 326 333 PF00069 0.792
MOD_GSK3_1 336 343 PF00069 0.669
MOD_GSK3_1 358 365 PF00069 0.711
MOD_GSK3_1 472 479 PF00069 0.586
MOD_GSK3_1 49 56 PF00069 0.821
MOD_GSK3_1 502 509 PF00069 0.740
MOD_GSK3_1 65 72 PF00069 0.576
MOD_GSK3_1 808 815 PF00069 0.802
MOD_GSK3_1 859 866 PF00069 0.792
MOD_GSK3_1 872 879 PF00069 0.556
MOD_GSK3_1 882 889 PF00069 0.589
MOD_GSK3_1 936 943 PF00069 0.653
MOD_GSK3_1 961 968 PF00069 0.675
MOD_N-GLC_1 133 138 PF02516 0.622
MOD_N-GLC_1 384 389 PF02516 0.532
MOD_N-GLC_1 812 817 PF02516 0.632
MOD_NEK2_1 1148 1153 PF00069 0.716
MOD_NEK2_1 1201 1206 PF00069 0.620
MOD_NEK2_1 328 333 PF00069 0.709
MOD_NEK2_1 367 372 PF00069 0.534
MOD_NEK2_1 427 432 PF00069 0.411
MOD_NEK2_1 670 675 PF00069 0.480
MOD_NEK2_1 956 961 PF00069 0.623
MOD_NEK2_2 1109 1114 PF00069 0.620
MOD_NEK2_2 506 511 PF00069 0.666
MOD_PIKK_1 344 350 PF00454 0.584
MOD_PIKK_1 936 942 PF00454 0.608
MOD_PK_1 708 714 PF00069 0.512
MOD_PKA_2 1028 1034 PF00069 0.659
MOD_PKA_2 1089 1095 PF00069 0.658
MOD_PKA_2 1117 1123 PF00069 0.672
MOD_PKA_2 153 159 PF00069 0.775
MOD_PKA_2 798 804 PF00069 0.700
MOD_Plk_1 1176 1182 PF00069 0.801
MOD_Plk_1 384 390 PF00069 0.525
MOD_Plk_1 859 865 PF00069 0.790
MOD_Plk_1 94 100 PF00069 0.664
MOD_Plk_2-3 882 888 PF00069 0.619
MOD_Plk_2-3 95 101 PF00069 0.795
MOD_Plk_4 1176 1182 PF00069 0.799
MOD_Plk_4 1223 1229 PF00069 0.627
MOD_Plk_4 278 284 PF00069 0.611
MOD_Plk_4 367 373 PF00069 0.527
MOD_Plk_4 384 390 PF00069 0.525
MOD_Plk_4 506 512 PF00069 0.664
MOD_Plk_4 708 714 PF00069 0.411
MOD_Plk_4 859 865 PF00069 0.785
MOD_Plk_4 876 882 PF00069 0.495
MOD_ProDKin_1 1071 1077 PF00069 0.691
MOD_ProDKin_1 1150 1156 PF00069 0.723
MOD_ProDKin_1 197 203 PF00069 0.731
MOD_ProDKin_1 358 364 PF00069 0.720
MOD_ProDKin_1 594 600 PF00069 0.512
MOD_ProDKin_1 908 914 PF00069 0.750
MOD_ProDKin_1 983 989 PF00069 0.512
MOD_SUMO_for_1 298 301 PF00179 0.734
MOD_SUMO_rev_2 1054 1063 PF00179 0.603
TRG_DiLeu_BaEn_1 422 427 PF01217 0.406
TRG_DiLeu_BaEn_1 844 849 PF01217 0.704
TRG_DiLeu_BaLyEn_6 1140 1145 PF01217 0.646
TRG_DiLeu_BaLyEn_6 596 601 PF01217 0.450
TRG_DiLeu_BaLyEn_6 923 928 PF01217 0.628
TRG_ENDOCYTIC_2 1231 1234 PF00928 0.733
TRG_ENDOCYTIC_2 1240 1243 PF00928 0.411
TRG_ENDOCYTIC_2 374 377 PF00928 0.515
TRG_ENDOCYTIC_2 428 431 PF00928 0.411
TRG_ENDOCYTIC_2 550 553 PF00928 0.411
TRG_ENDOCYTIC_2 557 560 PF00928 0.411
TRG_ENDOCYTIC_2 845 848 PF00928 0.631
TRG_ENDOCYTIC_2 905 908 PF00928 0.671
TRG_ENDOCYTIC_2 953 956 PF00928 0.713
TRG_ENDOCYTIC_2 974 977 PF00928 0.411
TRG_ENDOCYTIC_2 999 1002 PF00928 0.411
TRG_ER_diArg_1 526 528 PF00400 0.664
TRG_ER_diArg_1 604 606 PF00400 0.512
TRG_ER_diArg_1 700 702 PF00400 0.512
TRG_ER_diArg_1 805 808 PF00400 0.627
TRG_ER_diArg_1 837 840 PF00400 0.768
TRG_NES_CRM1_1 543 556 PF08389 0.512
TRG_NES_CRM1_1 603 617 PF08389 0.512
TRG_NLS_MonoExtC_3 522 527 PF00514 0.680
TRG_Pf-PMV_PEXEL_1 1143 1147 PF00026 0.661
TRG_Pf-PMV_PEXEL_1 700 704 PF00026 0.428

Homologs

Protein Taxonomy Sequence identity Coverage
A0A0N0P405 Leptomonas seymouri 48% 88%
A0A3S5H5Y6 Leishmania donovani 92% 77%
A4H509 Leishmania braziliensis 70% 83%
A4HT83 Leishmania infantum 94% 100%
E9AL71 Leishmania mexicana (strain MHOM/GT/2001/U1103) 84% 79%

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LeishMANIAdb - Server version: v0.0.2. - Database version: v0.0.1. - ChangeLog - © 2022-2024 Protein Bioinformatics Research Group, Institute of Enzymology, RCNS